As your primary observations are counts, you should use an analysis method that is designed for counts and thus models observations with 0 counts directly. A Poisson generalized linear model with log link is a standard way to start.
As Kjetil B Halvorsen said in a comment, you use an offset, log(area)
, in your regression to account for the area corresponding to each observation. Then you can interpret your results of modeling counts in the units of individuals/1000 hectares that you desire. See this page among many others on this site.
It's possible that a simple Poisson model won't be adequate, in that the equality between variance and mean for a Poisson distribution won't adequately describe your results. A "quasi-Poisson" model, in which confidence intervals are based on other-than-Poisson variance, or a negative binomial model could be next steps in that situation.
It's not clear from your description that a mixed model ("MM" in your abbreviations) would be needed here. Whether you need to consider a "generalized additive" ("GA") model has to do with whether simple functional forms of your covariates are adequate in the linear predictor of log(counts)
. I'd start with simple linear modeling of covariates in a Poisson generalized linear model, as if that works it's usually easier to explain to others.