Generally IncNodePurity is taken as main indicator of importance. From ?importance:
> set.seed(4543)
> data(mtcars)
> mtcars.rf <- randomForest(mpg ~ ., data=mtcars, ntree=1000,
keep.forest=FALSE, importance=TRUE)
>
> importance(mtcars.rf)
%IncMSE IncNodePurity
cyl 16.168645 169.96741
disp 18.672188 260.08722
hp 17.584375 184.95007
drat 6.948743 63.54528
wt 17.818509 254.30347
qsec 4.772889 33.25546
vs 5.303058 24.39064
am 5.210181 17.36626
gear 4.619161 21.55450
carb 8.577037 28.46715

Relative importance can be relative to minimum value of IncNodePurity:
X.IncMSE IncNodePurity relativeIncNodePurity
am 5.210181 17.36626 1.000000
gear 4.619161 21.55450 1.241171
vs 5.303058 24.39064 1.404484
carb 8.577037 28.46715 1.639221
qsec 4.772889 33.25546 1.914947
drat 6.948743 63.54528 3.659123
cyl 16.168645 169.96741 9.787219
hp 17.584375 184.95007 10.649965
wt 17.818509 254.30347 14.643536
disp 18.672188 260.08722 14.976581